Aquatic Salamanders - Found in water and on land
|
Adults of these species live mostly on land where they are active at night, but they also move to water during the breeding season where they lay eggs in water that hatch into small gilled fish-like larvae. The larvae live swimming in the water until they lose their gills then move onto the land to live, breathing with lungs and looking like miniature versions of the adults. Salamander larvae can be distinguished from tadpoles by the presence of gills, which are not found on frog larvae.
Occasionally, juveniles do not transform to live on land. They spend their lives living in water, breathing with gills. These neotenic adults (or paedomorphs) look like very large aquatic larvae.
Males and females of all of these species are similar enough in appearance that knowing the sex of a salamander will not help to identify a species.
|
Mole Salamanders - genus Ambystoma
|
Mole Salamanders live mostly underground except during rainy periods when they come above ground at night to breed and lay eggs in pools of water.
Juvenile Mole Salamanders hatch and live in pools of water until they lose their gills and move onto land.
|
California Tiger Salamander
Ambystoma californiense |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult © Jon Hirt |
Adult © Spencer Riffle |
Larvae are aquatic.
Gills are long - reduce in size with age. Dorsal fin is on back and tail. |
Historically found only in
the red areas |
|
|
|
|
|
|
|
|
Northwestern Salamander
Ambystoma gracile |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Larvae are aquatic.
Gills are long - reduce in size with age. Dorsal fin is on back and tail. |
Found only in the red area |
|
|
|
|
|
|
|
|
Long-toed Salamanders - Ambystoma macrodactylum
|
An elongated toe
can help to identify
Long-toed Salamanders |
|
|
|
|
|
|
|
The fourth toe on each back foot is longer than third toe. This is not found on other salamander species in California. |
© Spencer Riffle |
|
|
|
|
|
|
|
|
|
|
|
|
|
Santa Cruz
Long-toed Salamander
Ambystoma macrodactylum croceum |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult © Brad Alexander |
Juvenile © Jon Hirt |
Larvae are aquatic.
Gills are long - reduce in size with age. Dorsal fin is on back and tail. |
Found in the dark blue area
(Near Monterey Bay) |
|
|
|
|
|
|
|
|
Southern
Long-toed Salamander
Ambystoma macrodactylum sigillatum |
|
|
|
|
|
|
|
|
Adult |
Adult © Chad Lane |
Adult © William Flaxington |
Juvenile |
Larvae are aquatic.
Gills are long - reduce in size with age. Dorsal fin is on back and tail.
|
Found only in the red areas |
|
|
|
|
|
|
|
|
Western Tiger Salamanders are Mole Salamanders that are not native to California. They have been introduced into some Calfiornia lakes and ponds. Individuals found in California are probably derived from several suspecies found across the west, which accounts for
the various colors and patterns which can be found. Hybrids between Western Tiger Salamanders
and California Tiger Salamanders are also sometimes found in areas where California Tiger Salamanders occur.
|
|
|
|
|
|
|
|
|
Western Tiger Salamander
Ambystoma mavortium
|
|
|
|
|
|
|
|
Adult, Mono County © Adam Clause |
Adult |
Adult, Siskiyou County © Alan Barron |
Adult, Siskiyou County © Alan Barron |
Larvae are aquatic.
Gills are long - reduce in size with age. Dorsal fin is on back and tail. |
|
|
|
|
|
|
|
Adult, Kern County © William Flaxington |
Adult, San Diego County © Sean Kelly |
Adult, Lake County |
Hybrid with California Tiger Salamander |
Adult, Arizona |
Adult, Arizona |
Found in the red areas
and possibly elsewhere |
|
|
|
|
|
|
|
|
Giant Salamanders - genus Dicamptodon
|
There are two species of Giant Salamanders in California. They both look alike, so use the location to help identify the species. The Coastal Giant Salamander is found north of the Mendocino County line, the California Giant Salamander is south.
Giant Salamanders lay eggs in water that hatch into gilled larvae which live at the edge of streams until they lose their gills and move onto land. Aquatic neotenic adults are not uncommon. Adults are occasionally found walking near streams in daylight.
|
California
Giant Salamander
Dicamptodon ensatus |
|
|
|
|
|
|
|
|
Adult |
Juvenile |
Adult |
Juvenile |
Larvae are aquatic.
Gills are short. Dorsal fin is on tail, not back. |
Found only in the red areas |
|
|
|
|
|
|
|
|
Coastal
Giant Salamander
Dicamptodon tenebrosus
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Aquatic neotenic adult |
Larvae are aquatic.
Gills are short. Dorsal fin is on tail, not back. |
Found only in the red area |
|
|
|
|
|
|
|
|
Torrent Salamanders - genus Rhyacotriton
|
Like the aquatic salamanders Torrent Salamanders have very small lungs and are often found immersed in streams, but they are also similar to terrestrial salamanders in that they spend time out of the water where they need to remain moist. Eggs are laid in streams where they hatch into gilled larvae that live in the water until they lose their gills and move onto land.
|
Southern
Torrent Salamander
Rhyacotriton variegatus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult, © Zach Lim |
Larvae are aquatic.
Gills are very small. Dorsal fin is on tail, not on back |
Found only in the red area |
|
|
|
|
|
|
|
|
Pacific Newts - genus Taricha
|
Pacific Newts are the salamanders that are most likely to be seen walking around in daylight. They live under surface objects on land or underground but they emerge to move to water in Winter or Spring where they change into an aquatic phase that helps them swim. They are often seen walking or in ponds during their breeding migrations. They breed and lay eggs, then leave the water to live on land where they change back into their terrestrial phase. The eggs hatch into tiny gilled larva which eventually transform and leave the water to live on land.
Except for the Red-bellied Newt, the only species that has all-black eyes, the other three species can be hard to identify. Click on this link to find more information
to help with Pacific Newt identification, inluding larvae, in areas where more than one species occurs.
|
Rough-skinned Newt
Taricha granulosa
|
|
|
|
|
|
|
|
|
Terrestrial phase adult |
Terrestrial phase juvenile |
Terrestrial phase adult |
Aquatic phase adult |
Larvae are aquatic.
Stripes not present. Gills are long - reduce in size with age.
Dorsal fin is on back and tail. |
Found only in the red areas |
|
|
|
|
|
|
|
|
Red-bellied Newt
Taricha rivularis
|
|
|
|
|
|
|
|
|
Terrestrial phase adult |
Unlike the other Pacific Newts which have a yellow patch in their eyes, the eyes of Red-bellied Newts are all black. |
Aquatic phase adult |
Aquatic phase adult |
Larvae are aquatic.
Stripes not present. Gills are long - reduce in size with age.
Dorsal fin is on back and tail. |
Found only in the red area |
|
|
|
|
|
|
|
|
Sierra Newt
Taricha sierrae
|
|
|
|
|
|
|
|
|
Terrestrial phase adult |
Adult |
Terrestrial phase adult |
Aquatic phase adult |
Larvae are aquatic.
Stripes are present. Gills are long - reduce in size with age.
Dorsal fin is on back and tail. |
Found only in the orange area |
|
|
|
|
|
|
|
|
Coast Range Newt
Taricha torosa
|
|
|
|
|
|
|
|
|
Terrestrial phase adult |
Terrestrial phase adult |
Terrestrial phase adult |
Aquatic phase adult |
Larvae are aquatic. (left) young (right) mature
Stripes are present. Gills are long - reduce in size with age.
Dorsal fin is on back and tail. |
Found only in the red areas |
|
|
|
|
|
|
|
|
Terrestrial Salamanders - Family Plethodontidae (Found on land, not in water)
|
The following terrestrial salamanders all live in moist places on land, emerging at night to hunt. They are rarely found active in the daytime. They are typically found under rocks, logs, and other surface objects and they are occasionally seen walking at night. They need to stay moist but they are rarely found in water since they respire through their skin instead of using lungs. They breed and lay their eggs in moist places on land, not in water like the aquatic salamanders. The young hatch fully-formed as a miniature version of the terrestrial adults, but they are sometimes colored differently than adults. Males and females of all of these species are similar enough in appearance that knowing the sex of a salamander will not help to identify a species. |
|
Climbing Salamanders - genus Aneides
|
Climbing Salamanders have thick toe tips that are specially adapted to help them climb steep surfaces such as rocks and trees.
|
|
Black Salamanders - Aneides flavipunctatus complex - Found in several colors and patterns. There are four separate species but they do not correspond exactly to the different colors and patterns.
Hatchling juveniles often look very different from the adults with yellow or greenish coloring.
|
Speckled
Black Salamander
Aneides flavipunctatus
(Spotted Form)
|
|
|
|
|
|
|
|
|
Adult |
Sub-adult |
Adult © Zach Lim |
Adult and juvenile |
Juvenile |
Black Salamanders have
rounded toe tips
|
The Speckled Black Salamander
is found in the Purple area |
(Black Form) |
|
|
|
|
|
|
|
|
Adult |
Adult |
Juvenile |
|
|
|
|
(Frosted Form) |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Sub-adult |
Juvenile |
Juvenile |
|
|
|
|
|
|
|
|
|
Shasta
Black Salamander
Aneides iecanus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult © Jonathan Hakim |
Adult © Zach Lim |
Adult © Jonathan Hakim |
Juvenile © Mark Gary |
Found in the Orange areas |
|
|
|
|
|
|
|
|
Klamath
Black Salamander
Aneides klamathensis |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult © Alan Barron |
Adult |
Sub-adult © Alan Barron |
Juvenile © Alan Barron |
Found in the Red area |
|
|
|
|
|
|
|
|
Santa Cruz
Black Salamander
Aneides niger
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult © Spencer Riffle |
Adult and juvenile |
Juvenile © Zachary Lim |
Found in the Dark Blue area |
|
|
|
|
|
|
|
|
Arboreal Salamanders can have varying amounts of spotting on the body - from very fine spots to very large ones - and varying background colors, but they are all the same species.
Hatchling and very young juvenile Arboreal Salamanders do not look like the adults, having a very dark background with a wash of light markings, not just spots. |
Arboreal Salamander
Aneides lugubris
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Juvenile |
Hatchling © Spencer Riffle |
Found only in the red areas |
|
|
|
|
|
|
|
|
Clouded Salamanders are very similar to Wandering Salamanders in appearance and behavior, so use the range maps to help identify them where both species occur in California. Both species are typically found under the bark of fallen trees.
|
Clouded Salamander
Aneides ferreus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult © William Flaxington |
Sub-adult |
Juvenile |
Clouded Salamanders have
squared-lff toe tips |
Found only in the red area |
|
|
|
|
|
|
|
|
Wandering Salamander
Aneides vagrans |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Sub-adult |
Juvenile © Alan Barron |
Wandering Salamanders have
squared-off toe tips. |
Found only in the red area |
|
|
|
|
|
|
|
|
Slender Salamanders - genus Batrachoseps
|
Slender Salamanders are small worm-like salamanders with tiny legs that are sometimes not noticed.
The different species are hard to identify by appearance, so use the range maps to help identify them since in most areas only one species is found.
|
Range map of all Slender Salamanders in California:
|
|
|
|
|
|
|
|
|
|
Physical characteristics that
can help to identify
Slender Salamanders |
|
|
|
|
|
|
Slender Salmanders have only 4 toes on their hind feet.
All other California salamanders have
5 toes on their hind feet. |
Slender salamanders have long thin bodies. The tail can be longer that the body but the tail can also be broken off. The legs are very small and they are sometimes not seen. Slender Salamanders are sometimes mistaken for a worm or a tiny snake when the legs are not seen. |
In some areas of southern California two species of Slender Salamanders can be found. They can usually be identified by looking at the underside.
Left: The Garden Slender Salamander has a light underside.
Right:
The Black-bellied Slender Salamander has a dark underside. |
The Island Slender Salamander also has a light underside. It is found on Santa Cruz Island along with the Black-bellied Slender Salamander (left). |
|
Greenhorn Mountains
Slender Salamander
Batrachoseps altasierrae |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult © Ryan Sikola |
Adult |
Found only in the red area |
|
|
|
|
|
|
|
|
California
Slender Salamander
Batrachoseps attenuatus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Tiny Juvenile |
Found only in the red areas |
|
|
|
|
|
|
|
|
Fairview
Slender Salamander
Batrachoseps bramei |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Adult |
Found only in the red area |
|
|
|
|
|
|
|
|
Inyo Mountains
Slender Salamander
Batrachoseps campi |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult © Adam Clause |
Adult © Adam Clause |
Adult and juvenile, © Ryan Sikola |
Found only in the red areas |
|
|
|
|
|
|
|
|
Hell Hollow
Slender Salamander
Batrachoseps diabolicus
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Adult © Chad M. Lane |
Found only in the red area |
|
|
|
|
|
|
|
|
San Gabriel Mountains
Slender Salamander
Batrachoseps gabrieli
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Adult and Juvenile © Adam Clause |
Found only in the red areas |
|
|
|
|
|
|
|
|
Gabilan Mountains
Slender Salamander
Batrachoseps gavilanensis |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile |
Found only in the red area |
|
|
|
|
|
|
|
|
Gregarious
Slender Salamander
Batrachoseps gregarius |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Adult |
Found only in the red area |
|
|
|
|
|
|
|
|
San Simeon
Slender Salamander
Batrachoseps incognitus
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult © Spencer Riffle |
Adult © Ryan Sikola |
Adult © Spencer Riffle |
Juvenile © Ryan Sikola |
Found only in the red area |
|
|
|
|
|
|
|
|
Sequoia
Slender Salamander
Batrachoseps kawia |
|
|
|
|
|
|
|
|
Adult |
Adult
|
Adult © Brad Alexander |
Adult |
Adult |
Adult |
Found only in the red area |
|
|
|
|
|
|
|
|
Santa Lucia Mountains
Slender Salamander
Batrachoseps luciae
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult © Ryan Sikola |
Juvenile |
Found only in the red area |
|
Southern California Slender Salamander - Batrachoseps major |
Desert
Slender Salamander
Batrachoseps major aridus |
|
|
|
|
|
|
|
|
Adult © Mario Garcia-Paris |
Adult © Mario Garcia-Paris |
Adult © Arden Brame
(courtesy of Harold De Lisle.) |
|
|
|
Found in the bright blue area
(Isolated desert canyons in Riverside County only.)
|
|
|
|
|
|
|
|
|
Garden
Slender Salamander
Batrachoseps major major
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
The underside is light in color. |
Adult and juvenile |
Found in the red and blue areas |
|
|
|
|
|
|
|
|
|
Lesser
Slender Salamander
Batrachoseps minor |
|
|
|
|
|
|
|
|
Adult |
Adult © Spencer Williams |
Adult © Spencer Williams |
Adult © David Wake |
Adult © Ryan Sikola |
Adult © Spencer Riffle |
Found only in the red area |
|
|
|
|
|
|
|
|
Black-bellied
Slender Salamander
Batrachoseps nigriventris |
|
|
|
|
|
|
|
Adult |
Adult |
Adult © Ryan Sikola |
Adult |
The underside is dark in color. |
Adult and Juvenile |
Found only in the red area |
|
|
|
|
|
|
|
|
Channel Islands
Slender Salamander
Batrachoseps pacificus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult © Spencer Williams |
Adult |
The underside is light in color |
Found only in the red areas
(Channel Islands only) |
|
|
|
|
|
|
|
|
Kings River
Slender Salamander
Batrachoseps regius
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Adult |
Found only in the red area |
|
|
|
|
|
|
|
|
Relictual
Slender Salamander
Batrachoseps relictus |
|
|
|
|
|
|
|
|
Adult © William Flaxington |
Adult © Robb Schell |
Adult © Brad Alexander |
Adult © Rob Schell |
Adult © Mark Gary |
Adult and juveniles © Noah Morales |
Found only in the red area |
|
|
|
|
|
|
|
|
Kern Plateau
Slender Salamander
Batrachoseps robustus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult © Noah Morales |
Adult |
Found only in the red area |
|
|
|
|
|
|
|
|
Kern Canyon
Slender Salamander
Batrachoseps simatus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult © Ryan Sikola |
Adult |
Hatchling on U.S. ten-cent coin |
Found only in the red area |
|
|
|
|
|
|
|
|
Tehachapi
Slender Salamander
Batrachoseps stebbinsi
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Adult © Ryan Sikola
|
Found only in the red area |
|
|
|
|
|
|
Arguello
Slender Salamander
Batrachoseps wakei
|
|
|
|
|
|
|
Adult © Robert Hansen |
Adult © Robert Hansen |
|
|
Found only in the tiny red area |
|
|
|
|
|
|
|
|
Ensatina - Ensatina eschscholtzii
|
The genus Ensatina currently consists of one species - eschscholtzii - with seven different subspecies which can be identified by color and pattern and by range. (Ensatina klauberi is also recognized as a full species by some herpetologists.) |
Physical characteristics that can help to identify Ensatina |
|
|
|
|
|
|
|
Unlike other salamanders, the tail of an Ensatina is constricted at its base |
When threatened, Ensatina often assume a defensive stance with the legs and tails raised as they excrete a milky substance from their tail and neck. |
Juveniles of some subspecies are similar to adults, but other subspecies juveniles can be very dark, unlike adults, with orange or yellow on the legs and sometimes the tail |
Juveniles © Alan Barron |
Yellow-blotched Ensatina
Ensatina eschscholtzii
croceater |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile |
Found in the dark blue areas |
|
|
|
|
|
|
|
|
Monterey Ensatina
Ensatina eschscholtzii eschscholtzii |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile |
Found only in the purple area |
|
|
|
|
|
|
|
|
Large-blotched Ensatina
Ensatina eschscholtzii
klauberi |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile |
Found in the bright blue area |
|
|
|
|
|
|
|
|
Oregon Ensatina
Ensatina eschscholtzii oregonensis
(California intergrades)
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile |
Found only in the red area |
|
|
|
|
|
|
|
|
Painted Ensatina
Ensatina eschscholtzii
picta |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Juveniles © Alan Barron |
Juvenile |
Found only in the pink area |
|
|
|
|
|
|
|
|
Sierra Nevada Ensatina
Ensatina eschscholtzii
platensis |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult and juvenile |
Juvenile |
Found only in the orange area |
|
|
|
|
|
|
|
|
Yellow-eyed Ensatina
Ensatina eschscholtzii
xanthoptica |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult without tail |
Juvenile |
Found only in the yellow areas |
|
|
|
|
|
|
|
|
Web-footed Salamanders - genus Hydromantes
|
Web-footed Salamanders are associated with moist rocky areas, including granite and limestone cliffs, caves, and caverns. The rear feet have slightly-webbed toes which aid in climbing steep rock surfaces. They can be easily identified by their location, but in the area around Lake Shasta there are three similar species, so be sure to check the enlarged range map.
|
Physical characteristics that can help to identify
Web-footed Salamanders |
|
|
|
|
|
|
|
The rear feet have webbed toes |
Webbed toes |
Webbed toes |
The body can be flattened to
fit into narrow rock crevices |
Adult in rock crack © Spencer Williams |
|
|
Limestone Salamander
Hydromantes brunus
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juveniles are greenish yellow |
Found only in the red area |
|
|
|
|
|
|
|
|
Mount Lyell Salamander
Hydromantes platycephalus
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile |
Found only in the red area |
|
|
|
|
|
|
|
|
Samwel Shasta Salamander
Hydromantes samweli |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult and juvenile |
Juvenile |
Found only in the yellow area
See enlarged range map
|
|
|
|
|
|
|
|
Shasta Salamander
Hydromantes shastae |
|
|
|
|
|
|
|
Adult © Noah Morales |
Adult © Noah Morales |
Adult © Spencer Riffle |
Adult © Spencer Riffle |
Adult © Noah Morales |
Found only in the orange area
See enlarged range map
|
|
|
|
|
|
|
|
|
Wintu Shasta Salamader
Hydromantes wintu |
|
|
|
|
|
|
|
|
Adult © Spencer Williams |
Adult © Noah Morales |
Adult © Noah Morales |
Adult © Noah Morales |
Adult © Spencer Williams |
Juvenile © Noah Morales |
Found only in the red area
See enlarged range map |
|
Woodland Salamanders - genus Plethodon
|
Woodland Salamanders found in California are similar in size and appearance, but their ranges are limited where they are only found in woodlands in the extreme northwest part of the state. Use the range maps to help identify them.
|
Scott Bar Salamander
Plethodon asupak |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult © Alan Barron |
Adult, and juvenile © Tim Burkhardt
|
Juveniles have reddish
stripes on the back |
Found only in the red area |
|
|
|
|
|
|
|
|
Dunn's Salamander
Plethodon dunni
|
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult |
Juvenile © Alan Barron |
Found only in the red area |
|
|
|
|
|
|
|
|
Del Norte Salamander
Plethodon elongatus |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult and juvenile |
Juveniles |
Found only in the red area |
|
|
|
|
|
|
|
|
Siskiyou Mountains
Salamander
Plethodon stormi |
|
|
|
|
|
|
|
|
Adult |
Adult |
Adult |
Adult |
Adult © Spencer Riffle |
Juvenile |
Found only in the red area |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|