CaliforniaHerps.com

A Guide to the Amphibians
and Reptiles of California




Island Night Lizard - Xantusia riversiana

San Clemente Night Lizard - Xantusia riversiana reticulata

Smith, 1946
Click on a picture for a larger view
Island Night Lizards Range Map
Red: Range of this subspecies
Xantusia riversiana reticulata - San Clemente Night Lizard

Bright Green: Range of the other subspecies
Xantusia riversiana riversiana - San Nicolas Night Lizard


Click on the map for a topographical view

Map with California County Names




observation link





San Clemente Night Lizard
Adult, Santa Barbara Island
San Clemente Night Lizard San Clemente Night Lizard San Clemente Night Lizard
Adult, Santa Barbara Island Adult, Santa Barbara Island
San Clemente Night Lizard San Clemente Night Lizard San Clemente Night Lizard
Adult, Santa Barbara Island Adult, Santa Barbara Island Adult, Santa Barbara Island
San Clemente Night Lizard San Clemente Night Lizard San Clemente Night Lizard
Adult, Santa Barbara Island Adult, Santa Barbara Island Adult, Santa Barbara Island
San Clemente Night Lizard San Clemente Night Lizard San Clemente Night Lizard
Adult, Santa Barbara Island Adult, Santa Barbara Island Adult, Santa Barbara Island
San Clemente Night Lizard San Clemente Night Lizard San Clemente Night Lizard
Adult, striped form, San Clemente Island
© Bob Haase
Adult, San Clemente Island
© Bob Haase
Adult, San Clemente Island
© Bob Haase
  Night lizard scales  
  Island Night Lizards have small granular scales on soft skin.  
     
Habitat
San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat
Santa Barbara Island Santa Barbara Island Santa Barbara Island and Sutil Islet,
early Summer 
San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat
Santa Barbara Island,
early Summer 
Grassy plateau, top of
Santa Barbara Island
View looking down at Elephant Seal Beach, Santa Barbara Island
San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat
Rocky cliff-edge and Sutil Islet, early Summer, Santa Barbara Island
Dormant Giant Coreopsis, Santa Barbara Island, early summer
San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat
Santa Barbara Island, early Summer  Santa Barbara Island landing area Santa Barbara Island, early Summer 
San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat San Clemente Night Lizard Habitat
Santa Barbara Island with Spring greenery and blooming Giant Coreopsis  © Tim Burkhardt
   
Description
 
Size
2.5 - 4.2 inches long from snout to vent (6.3 - 10.6 cm). (Stebbins 2003)
The largest of the Xantusiids, growing from about 5 - 8 inches long including the tail.

Appearance
A medium-sized lizard with granular scales, soft skin that appears loose around the neck and shoulders, large plates on the head, lidless eyes with vertical pupils, a gular fold, and a fold of skin low on each side of the body.
Two rows of supraoculars and 16 lengthwise rows of squarish scales at the midbelly.
Color and Pattern
Color and pattern are variable - Brown, olive-brown, grayish, rusty, or yellowish-brown with dark brown or black spots or blotches, sometimes with full or broken light stripes edged with black along the sides of the back.
Striped lizards occur on San Clemente Island, but rarely on Santa Barbara Island.
The underside is slate bluish, cream or pale yellow.
The undersides of the feet are sometimes yellowish.

Life History and Behavior

Activity
Once considered nocturnal, but apparently diurnal but sedentary, secretive, and rarely seen.
Spends most of its time under cover, but occasionally seen exposed on the surface in daylight.
The mild climate of the islands this lizard inhabits allows it to remain active throughout the year.
Grows and matures slowly, living to at least 12 years old
Defense
The tail can break off easily, but it will grow back.
The detached tail wriggles on the ground which can distract a predator from the body of the lizard allowing it time to escape.
More information about tail loss and regeneration.
Diet and Feeding
Eats small invertebrates including insects, spiders, scorpions, and marine isopods along with plant material (which can make up to one-third of its diet.)
Reproduction
Reproductive potential is low.
Adults do not reach sexual maturity until their 3rd or 4th year and only about half of all females reproduce each year.
Breeds in March and April.
Viviporous, bearing 2-9 live young mostly in September.

Habitat
Found in almost any island habitat that provides it protection and shade - maritime desert scrub, grassland, chaparral, oak savanna, cactus, dry streambeds, cliffs, rocky beaches, sparsely-vegetated areas. Takes shelter in cracks in rocks or in the ground, and under surface objects such as rocks, fallen vegetation and beach driftwood.

Santa Barbara Island is a flat tree-less plateau with grasses, small shrubs, introduced ice plant, prickly-pear cactus and Giant Coreopsis. There are a few small rocky canyons along the edge with dense vegetation.

Geographical Range
Endemic to California.
Found only on islands off the southern California coast - Santa Barbara Island, Sutil Islet, and San Clemente Island.

Notes on Taxonomy
Two subspecies of X. riversiana are still recognized by on the official lists of the SSAR and the CNAH:
X. r. reticulata

X. r. riversiana

"Although not mentioned by Noonan et al. (2013, Mol. Phylogenet. Evol. 69: 109–122), their results support the taxonomic distinction between populations of X. riversiana on San Nicolas Island (X. r. riversiana) and those on San Clemente and Santa Barbara Islands (X. r. reticulata). "
(SSAR Herpetological Circular No. 43, 2017.)


Alternate and Previous Names (Synonyms)

Xantusia riversiana - Island Night Lizard (Stebbins 1985, 2003)
Klauberina riversiana - Island NIght Lizard (Stebbins 1966)
Xantusia riversiana - Island Night Lizard (Smith 1946, Stebbins 1954)
Xantusia riversiana - Island Night Lizard (Rivers's Lizard) (Grinnell and Camp 1917)
Conservation Issues  (Conservation Status)
Protected since 1967. This lizard's restricted habitat has been threatened by the grazing of introduced livestock and game, (most of which have now been removed) and by introduced predators such as feral cats and possibly rats. It is also preyed upon heavily by native birds and mammals. In 1990 the USFWS categorized the status of lizards on San Clemente Island as stable. A 1991 study by Fellers and Drost determined that the population on Santa Barbara Island is not threatened.
The U.S. Navy administers San Clemente and San Nicolas Islands, and has petitioned to have lizards from those islands removed from the Endangered Species List. Santa Barbara Island and Sutil Islet are protected as National Park property.

On 4/1/14 the U.S. Fish and Wildlife Service removed Xantusia riversiana, the island night lizard, from the Federal List of Endangered and Threatened Wildlife. More Information
Taxonomy
Family Xantusiidae Night Lizards Baird, 1858
Genus Xantusia Night Lizards Baird, 1859 “1858”
Species riversiana Island Night Lizard Cope, 1883
Subspecies

reticulata San Clemente Night Lizard Smith, 1946
Original Description
Xantusia riversiana - Cope, 1883 - Proc. Acad. Nat. Sci. Philadelphia, Vol. 35, p. 29
Xantusia riversiana reticulata - Smith, 1946 - Journ. Washington Acad. Sci., Vol. 36, p. 392

from Original Description Citations for the Reptiles and Amphibians of North America © Ellin Beltz

Meaning of the Scientific Name
Xantusia - honors Xantus, John
riversiana
- honors Rivers, James, J.
reticulata - Latin - net-like - probably refers to the pattern.

from Scientific and Common Names of the Reptiles and Amphibians of North America - Explained © Ellin Beltz

Related or Similar California Lizards
X. henshawi - Granite Night Lizard
X. gracilis - Sandstone Night Lizard
X. vigilis - Yucca Night Lizard
X. sierrae - Sierra Night Lizard
X. r. riversiana - San Nicolas Night Lizard

More Information and References
California Department of Fish and Wildlife

US Fish & Wildlife Service

IUCN Red List

Robert L. Bezy. Night Lizards: Field Memoirs and a Summary of the Xantusiidae. ECO Herpetological Publishing & Distribution. 2019.

Stebbins, Robert C., and McGinnis, Samuel M.  Field Guide to Amphibians and Reptiles of California: Revised Edition (California Natural History Guides) University of California Press, 2012.

Stebbins, Robert C. California Amphibians and Reptiles. The University of California Press, 1972.

Flaxington, William C. Amphibians and Reptiles of California: Field Observations, Distribution, and Natural History. Fieldnotes Press, Anaheim, California, 2021.

Samuel M. McGinnis and Robert C. Stebbins. Peterson Field Guide to Western Reptiles & Amphibians. 4th Edition. Houghton Mifflin Harcourt Publishing Company, 2018.

Stebbins, Robert C. A Field Guide to Western Reptiles and Amphibians. 3rd Edition. Houghton Mifflin Company, 2003.

Behler, John L., and F. Wayne King. The Audubon Society Field Guide to North American Reptiles and Amphibians. Alfred A. Knopf, 1992.

Powell, Robert., Joseph T. Collins, and Errol D. Hooper Jr. A Key to Amphibians and Reptiles of the Continental United States and Canada. The University Press of Kansas, 1998.

Thelander, C. G., ed. Life on the Edge: A Guide to California's Endangered Natural Resources Volume I: Wildlife.
Biosystems Books, Santa Cruz, California. 1994.

Schoenherr, Allan A. Natural History of the Islands of California. The University of California Press. 2003.

Fellers, G.M. and Drost, C.A. 1991. Xantusia riversiana. Catalogue of American Amphibians and Reptiles 516: 1-4.

U.S. Fish and Wildlife Service (USFWS). 1990. Endangered and threatened species recovery program: report to Congress.

Bartlett, R. D. & Patricia P. Bartlett. Guide and Reference to the Turtles and Lizards of Western North America (North of Mexico) and Hawaii. University Press of Florida, 2009.

Jones, Lawrence, Rob Lovich, editors. Lizards of the American Southwest: A Photographic Field Guide. Rio Nuevo Publishers, 2009.

Smith, Hobart M. Handbook of Lizards, Lizards of the United States and of Canada. Cornell University Press, 1946.

Joseph Grinnell and Charles Lewis Camp. A Distributional List of the Amphibians and Reptiles of California. University of California Publications in Zoology Vol. 17, No. 10, pp. 127-208. July 11, 1917.

Conservation Status

The following conservation status listings for this animal are taken from the January 2024 State of California Special Animals List and the January 2024 Federally Listed Endangered and Threatened Animals of California list (unless indicated otherwise below.) Both lists are produced by multiple agencies every year, and sometimes more than once per year, so the conservation status listing information found below might not be from the most recent lists. To make sure you are seeing the most recent listings, go to this California Department of Fish and Wildlife web page where you can search for and download both lists:
https://www.wildlife.ca.gov/Data/CNDDB/Plants-and-Animals.

A detailed explanation of the meaning of the status listing symbols can be found at the beginning of the two lists. For quick reference, I have included them on my Special Status Information page.

If no status is listed here, the animal is not included on either list. This most likely indicates that there are no serious conservation concerns for the animal. To find out more about an animal's status you can also go to the NatureServe and IUCN websites to check their rankings.

The July 2023 Special Animals List and Federally Listed Endangered and Threatened Animals of California list desinations apply to the full species, Xantusia riversiana - Island Night Lizard.

Organization Status Listing  Notes
NatureServe Global Ranking G3 Vulnerable—At moderate risk of extinction due to a restricted range, relatively few populations (often 80 or fewer), recent and widespread declines, or other factors.
NatureServe State Ranking S3 Vulnerable in the state due to a restricted range, relatively few populations (often 80 or fewer), recent and widespread declines, or other factors making it vulnerable to extirpation from the state.
U.S. Endangered Species Act (ESA) Recovered & Delisted FT - Listed as Threatened 9/12/1977
FPD - Federally Proposed (Delisting) 2/4/2013
Recovered - Removed from ESA list 5/1/2014
California Endangered Species Act (CESA) None
California Department of Fish and Wildlife None
Bureau of Land Management None
USDA Forest Service None
IUCN LC Least Concern


 

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